Abstract
Background
   Depression is widespread global problem that not only severely impacts
   individuals’ physical and mental health but also imposes a heavy
   disease burden on nations and societies. The role of inflammation in
   the pathogenesis and pathophysiology of depression has received much
   attention, but the precise relationship between the two remains
   unclear. This study aims to investigate the correlation between
   depression and inflammation using a network medicine approach.
Methods
   We utilized a degree-preserving approach to identify the large
   connected component (LCC) of all depression-related proteins in the
   human interactome. The LCC was deemed as the disease module for
   depression. To measure the association between depression and other
   diseases, we calculated the overlap between these disease protein
   modules using the Sab algorithm. A smaller Sab value indicates a
   stronger association between diseases. Building on the results of this
   analysis, we further explored the correlation between inflammation and
   depression by conducting enrichment and pathway analyses of critical
   targets. Finally, we used a network proximity approach to calculate
   drug-disease proximity to predict the efficacy of drugs for the
   treatment of depression. We calculated and ranked the distances between
   depression disease modules and 6,100 drugs. The top-ranked drugs were
   selected to explore their potential for treating depression based on
   the hypothesis that their antidepressant effects are related to
   reducing inflammation.
Results
   In the human interactome, all depression-related proteins are clustered
   into a large connected component (LCC) consisting of 202 proteins and
   multiple small subgraphs. This indicates that depression-related
   proteins tend to form clusters within the same network. We used the 202
   LCC proteins as the key disease module for depression. Next, we
   investigated the potential relationships between depression and 299
   other diseases. Our analysis identified over 18 diseases that exhibited
   significant overlap with the depression module. Where S[AB] = −0.075
   for the vascular disease and depressive disorders module,
   S[AB] = −0.070 for the gastrointestinal disease and depressive
   disorders module, and S[AB] = −0.062 for the endocrine system disease
   and depressive disorders module. The distance between them S[AB] < 0
   implies that the pathogenesis of depression is likely to be related to
   the pathogenesis of its co-morbidities of depression and that potential
   therapeutic approaches may be derived from the disease treatment
   libraries of these co-morbidities. Further, considering that the
   inflammation is ubiquitous in some disease, we calculate the overlap
   between the collected inflammation module (236 proteins) and the
   depression module (202 proteins), finding that they are closely related
   (S[di] = −0.358) in the human protein interaction network. After
   enrichment and pathway analysis of key genes, we identified the HIF-1
   signaling pathway, PI3K-Akt signaling pathway, Th17 cell
   differentiation, hepatitis B, and inflammatory bowel disease as key to
   the inflammatory response in depression. Finally, we calculated the
   Z-score to determine the proximity of 6,100 drugs to the depression
   disease module. Among the top three drugs identified by drug-disease
   proximity analysis were Perphenazine, Clomipramine, and Amitriptyline,
   all of which had a greater number of targets in the network associated
   with the depression disease module. Notably, these drugs have been
   shown to exert both anti-inflammatory and antidepressant effects,
   suggesting that they may modulate depression through an
   anti-inflammatory mechanism. These findings demonstrate a correlation
   between depression and inflammation at the network medicine level,
   which has important implications for future elucidation of the etiology
   of depression and improved treatment outcomes.
Conclusion
   Neuroimmune signaling pathways play an important role in the
   pathogenesis of depression, and many classes of antidepressants
   exhibiting anti-inflammatory properties. The pathogenesis of depression
   is closely related to inflammation.
   Keywords: depression, inflammation, network medicine, medical
   treatments, gene
1. Introduction
   Depression is a severe mental system disorder that afflicts
   approximately 300 million people worldwide, and depression is
   characterized by high prevalence, high disability, and high suicide
   rates ([33]1). Depression imposes a heavy burden of illness on
   individuals and society, and it has become the leading cause of mental
   health-related disease burden worldwide ([34]2). There are few
   clinically available medications for depression, and the low cure rate,
   poor compliance, and high relapse rate have caused great distress to
   patients, their families, and society ([35]3). Antidepressant drugs are
   commonly employed in clinical settings as a primary means of treating
   depression. However, despite this approach, only about one-third of
   patients ultimately exhibit positive responses to these interventions.
   Furthermore, while such drugs do provide some measure of therapeutic
   efficacy, their high cost and significant risk for side effects present
   obstacles to broader application ([36]4). Therefore, there is an urgent
   need to discover new and better treatments. However, the etiology of
   depression remains a mystery, which is a severe obstacle to providing
   better clinical treatment options for depressed patients. Considerable
   human and material resources have been devoted to research on the
   etiology and treatment of depression. The role of neuroimmune in the
   pathogenesis and pathophysiology of depression has received much
   attention. Studies suggest that immune activation and cytokine
   production may be associated with depression and that cytokines are
   signaling proteins that mediate and regulate immune responses and
   inflammation. Many clinical studies have reported a correlation between
   depressive symptoms and concentrations of pro-inflammatory factor
   cells. Moreover, it has been shown that pharmacological
   anti-inflammatory interventions can successfully reduce both depressive
   symptoms and pro-inflammatory cytokine concentrations. This suggests a
   bidirectional causal relationship likely exists between inflammation
   and the onset of depressive symptoms ([37]5). Whether inflammatory
   markers can serve as biomarkers of depression still needs to be
   investigated in depth. Here, we utilized the ideas and methods of human
   proteome-based network medicine analysis in ([38]6) in the disease
   genetics and pharmacogenomics to explore the relationship between
   depression and inflammation. Compared with the previous researches
   about the depression and the inflammation, more relationships were
   introduced into our research, including disease-disease relation,
   disease-inflammation relation, drug-disease relation,
   drug-anti-inflammation relation and so on. Not only interpret the
   neuroimmunological mechanism of depression pathogenesis, but also
   provide hypotheses and ideas for treating and rationalizing depression.
   Overall, our studies have contributed to advancing the in-depth
   understanding of the relationship between neuroinflammation and
   depression. By utilizing techniques from networked medicine, we have
   simplified the complexity of traditional laboratories, improved
   research efficiency, and reduced research costs. These advantages have
   enabled researchers in the field of depression and inflammation to
   discover new treatments and innovative strategies more quickly. We
   trust that our study offers promising solutions for further development
   in this field.
2. Methods
2.1. Target screening for depression
   To get the targets relevant to depression, we searched the MeSH[39]^1
   database for medical keywords related to depression. Then we used the
   medical keyword in GeneCards[40]^2 and CTD[41]^3 databases
   Depression-related risk genes were retrieved. Because the Score value
   in the Genecards and CTD databases is an important indicator reflecting
   the correlation between the target and the disease, the target more
   closely related to depression is found based on the Score greater than
   the median as the screening criterion. Then 239 depression targets were
   screened to be standardized with the criterion and rule of the
   Uniprot[42]^4 database after duplicates were deleted. Finally, we
   localized these risk genes to the human protein interaction network and
   constructed a disease module for depression. Since different factors or
   pathways often contribute to the development of diseases, we utilized
   enrichment analysis to categorize the differential genes or substances
   based on their functions, in order to establish functional and
   phenotypic correlations.
2.2. Disease co-morbidity measurement
   The comorbidity of diseases often indicates the existence of shared
   molecular mechanisms in their etiology, and studying disease
   co-occurrence is crucial for accurate diagnosis, effective treatment,
   and better prognosis ([43]7). Therefore, before investigating the
   relationship between depression and inflammation, we first explored
   diseases that overlap with depression modules in the human protein
   interaction network (HPI network). The used HPI network comes from the
   literature ([44]6). Our aim was to examine the connection between
   depression comorbidities and inflammation by taking into account
   existing clinical studies. Here we use a corpus of all 299 diseases
   defined by the Medical Subject Headings (MeSH) ontology, collected and
   compiled by the author Barabási ([45]8) to explore the overlapping
   relationship between these 299 disease proteins and the depression
   disease module. We utilized the S[AB] (A is a sub-network of the
   depression disease proteins, B is one of other 299 diseases proteins)
   metric to evaluate the network-based overlap between the protein set of
   different diseases, where S[AB] < 0 signals a network-based overlap
   between the depression disease A and the gene associated with disease
   B. Next, S[AB] was defined as follows:
   [MATH: SAB=〈dAB〉−
   〈dAA〉+
   〈dBB〉2 :MATH]
   Where
   [MATH: 〈dAB〉 :MATH]
   presents the average of the shortest path distance between nodes in the
   depression targets A and nodes in the other disease targets B. The
   equation of
   [MATH: 〈dAB〉 :MATH]
   is:
   [MATH: dAB=1||B||
   ∑j∈B<
   /mi>mini
   ∈Adi,j,
   :MATH]
   where
   [MATH: d(i,j)
   :MATH]
   is the length value of the shortest path between protein i of
   depression disease and protein j of B disease in the human protein
   interaction network (HPI network).
   [MATH: ||B||
   :MATH]
   presents the number of proteins in B disease. Similarly, the definition
   of
   [MATH: 〈dAA〉 :MATH]
   is:
   [MATH: dAA=1||A||
   ∑j∈A<
   /mi>mini
   ∈Adi,j.
   :MATH]
2.3. Network-based enrichment and pathway analysis
   We performed enrichment and pathway analysis of the screened vital
   targets based on a network medicine approach to decipher the
   inflammatory mechanisms of depression. We constructed the depression
   disease module based on the human protein interaction network and used
   the algorithm of network proximity ([46]8) to explore the relationship
   between the disease module and inflammation critical targets in the
   network. The screened essential genes were then enriched to analyze the
   possible pathway of the neuroimmune mechanism of depression occurrence
   based on the important relevant disease ontology and pathways in the
   enrichment results. The functional annotation of the screened key
   targets could be obtained based on the String database.[47]^5 The
   screened key genes were further used to analyze the biological
   processes, molecular functions, cellular components, and signaling
   pathways. We expect to discover the new role of inflammatory pathways
   in the pathogenesis of depression. Essential cytokines may be
   identified to serve as biological markers for the diagnosis of
   depression. It will be helpful to provide new reference ideas and a
   basis for future clinical diagnosis of depression, and effectively
   predict the effect of antidepressant drug treatment. The better
   treatment plan promptly can be selected for the depressed patient based
   on his individualized characteristics.
2.4. Drug-disease proximity
   Network-based can be used as a valid tool to explore the link between
   different diseases and existed drugs and measure the effect of drug
   treatment ([48]9). Therefore, similar to the measurement in ([49]6,
   [50]9), drug-disease proximity of the top-ranked drugs were used to
   analyze their action mechanisms, which could reveal the
   anti-inflammatory effects of antidepressants in the clinic as well as
   the potential antidepressants from the existed drugs. Since the
   DrugBank database provided the drug-target information and drug
   indication information ([51]10), we employ the network proximity
   approach ([52]6, [53]9) to conduct the quantitative analysis between
   6,100 drugs targets collected by ([54]6) and the depression disease
   module. In this method, the Z-value metric is used to evaluate the
   proximity between a disease and a drug. The process of obtaining the
   Z-value first calculates the mean value of all the closet path lengths
   between drug targets and diseases to get the distance d[c].
   Specifically, given the set of depression proteins set A and the set of
   drug targets T, we calculate the distance
   [MATH:
   dc(A
   ,T)
   :MATH]
   by integrating the shortest path length between nodes s and t in the
   network, as follows:
   [MATH:
   dcA,T=
   1||T||∑t∈T<
   /mi>mina
   ∈Ada,t :MATH]
   However, the set sizes of various drugs are different, which leads to
   bias and unfair comparison, if we only use d[c]. as the proximity
   without other processes. Therefore, we created a reference distance
   distribution corresponding to the expected distances between two
   randomly selected groups of proteins matching the size and the degrees
   of the original depression proteins and drug targets in the network.
   The reference distance distribution was generated by calculating the
   distance between these randomly selected groups, a procedure repeated
   100 times. The mean
   [MATH:
   μdc
   (A,T) :MATH]
   and S.D.
   [MATH:
   δdc
   (A,T) :MATH]
   of the reference distribution were used to convert an observed distance
   to a normalized distance, defining the proximity measure:
   [MATH: Z(A,T)=
   dc(A,T)−
   μdc(
   A,T)δdc(A,T) :MATH]
   Obviously, the smaller the Z-value, the better the efficacy of the drug
   and the higher the ranking of the drug. To assess their expected
   efficacy against depression, we performed a ranking of these drugs and
   conducted the common analysis. We found that most of the FDA-approved
   antidepressants were ranked highly, presenting that calculating the
   drug-disease proximity (Z-value) has practical significance. More
   interestingly, we found that these top-ranked drugs often have
   anti-inflammatory effects in clinical treatment of corresponding
   diseases. This implies that anti-inflammatory strategies can alleviate
   depressive symptoms to some extent. In addition, some drugs with
   similar disease modules as depression rank high on the list. We
   speculate that they may have a positive effect on the treatment of
   depression, although more clinical trials are needed to determine the
   results.
3. Results
   The clinical diagnosis of depression is not only based on the patient’s
   clinical manifestations but also includes scale evaluation and relevant
   physical and chemical inspection ([55]11). However, for some patients
   with atypical symptoms of depression, specific diagnostic indicators
   may be lacking ([56]12). Therefore, it is important to accurately
   identify the pathogenesis of depression and to clarify the etiology of
   depression for both differential diagnosis and therapeutic care of
   depressed patients. A large body of clinical evidence suggests that
   neuroimmune plays an important role in the pathogenesis of depression
   and that the interaction of the central nervous system and inflammatory
   pathways are relevant to its development ([57]13, [58]14). In the
   medical field, the concept of inflammation is frequently used but its
   clinical meaning can be ambiguous and dependent on the specific
   clinical context ([59]15). One commonly employed strategy in drug
   development is “repurposing”—the process by which a known drug or
   treatment is applied to a new disease manifestation of interest. This
   approach enables rapid localization of potential therapeutic targets,
   substantially reducing clinical exploration time.
   To search for diseases that overlap with depression in the network, we
   first mapped the depression disease module (consisting of 239 host
   protein targets) onto a human protein interaction network comprising
   18,508 proteins and 332,749 physical interactions ([60]Figure 1). The
   nodes in the network represent genes or their corresponding gene
   products, the edges refer to the connection relationship between nodes
   and nodes, and the network constituted by continuous phases between
   nodes in the network is the maximum connected module. We analyzed the
   distribution of overlapping relationships between the depressive
   disorder module and 299 other disorders, as shown in [61]Figure 2. We
   found that among the relationships between disease modules and 299
   disease-related proteins, depression and cardiovascular diseases,
   gastrointestinal diseases, and endocrine diseases have significant
   overlapping relationships, such as coronary heart disease, inflammatory
   bowel disease, and diabetes mellitus, which often appear together with
   depression as comorbidities ([62]Table 1). These findings have been
   verified in numerous clinical studies demonstrating that patients with
   cardiovascular disease are more likely to experience depression than
   the general population, and that depressed patients have a higher risk
   of developing cardiovascular disease and mortality rate ([63]16).
   Inflammation plays a crucial role in the development and progression of
   cardiovascular disease, with inflammation acting as a trigger for the
   early stages of the atherosclerotic process. Patients with increased
   inflammatory cytokines are at an elevated risk of developing
   cardiovascular diseases (CVD). Additionally, numerous studies indicate
   that depression often co-occurs with gastrointestinal diseases such as
   functional dyspepsia (FD), inflammatory bowel disease (IBD), gastritis,
   gastric ulcer, and acute enteritis, with inflammatory cytokines
   exerting an important pathogenic role in the development of these
   disorders ([64]17). Similarly, a growing number of studies have
   demonstrated the correlation between depression and metabolic diseases.
   Depression often co-morbid with multiple metabolic diseases, including
   obesity and diabetes, and the underlying mechanisms of both
   pathogeneses involve chronic inflammation, which strongly correlates
   with disease severity. In summary, by exploring the overlap of
   depression modules with other disease proteins, this approach reveals
   the comorbid features of depression to some extent. By comparing these
   common comorbidities, we can speculate that they share similar
   pathogenesis. Thus, inflammation is likely to be their shared
   pathogenesis. This also implies that potential treatments for
   depression are likely to be derived from disease-specific treatments.
Figure 1.
   Figure 1
   [65]Open in a new tab
   Depression disease module. Depression-targeted proteins are not
   randomly distributed in the human interactome but form a large
   connectivity component (LCC) consisting of 202 proteins and multiple
   small subgraphs.
Figure 2.
   Figure 2
   [66]Open in a new tab
   Distribution of the network overlap measure S[AB] between 299 diseases
   and depression targets. S[AB] values represent the network-based
   overlap between depression targets A and the genes associated with each
   disease B.
Table 1.
   The top 10 where depression targets overlap with 299 other disease
   networks.
   Number Disease                   S[AB]
   1      Vascular diseases         −0.075
   2      Gastrointestinal diseases −0.070
   3      Cardiovascular diseases   −0.067
   4      Digestive system diseases −0.063
   5      Endocrine system diseases −0.062
   6      Neurologic manifestations −0.050
   7      Intestinal diseases       −0.045
   8      Neoplasms by site         −0.028
   9      Bone diseases             −0.027
   10     Heart diseases            −0.024
   [67]Open in a new tab
   Distance ranking of diseases that overlap with the depression disease
   module, where the smaller the S[AB], the larger the overlap between the
   two and the higher the ranking.
3.1. Analysis of key genes and pathways
   On top of the established disease modules, using this network overlap
   approach, we further explored the relationship between depression and
   inflammation. We screened immune-related genes and their products in
   peripheral blood and calculated the network relationship between them
   and the disease module. We found that the distance between them in the
   network S[d] = −0.358, i.e., the network formed by inflammation-related
   factors was included in the depression disease module, and the protein
   interaction network with 90 common targets was obtained by STRING
   database, with a total of 347 edges and a mean degree value of 7.71.
   The cytohubba tool in Cytoscape analyzed the PPI network, and the MCC
   method screened the top 10 key targets ([68]Table 2). Their network
   relationships are shown in [69]Figure 3. Among them, the
   pro-inflammatory factors TNF-α and IL-6 and the anti-inflammatory
   factors IL-4 and IL-10 are more central. And a large number of studies
   have shown that depressed patients with depression have C-reactive
   protein (CRP), prostaglandins, and other arachidonic acid derivatives,
   as well as IL-6, IL-1β and TNF-α were significantly increased ([70]18).
   In addition, some data suggest that serum cytokine concentrations,
   including IL-6, are associated with the severity and duration of
   depressive illness and the effectiveness of antidepressant medication
   ([71]19).
Table 2.
   The top 10 key targets were screened by the MCC method.
   Rank Name  Score
   1    IL6   43,964
   2    IL10  43,712
   3    TNF   42,358
   4    IL4   41,906
   5    IL1B  41,330
   6    IL18  40,442
   7    CXCL8 40,440
   8    IL1A  40,350
   9    CCL2  40,344
   10   IL17A 3,120
   [72]Open in a new tab
   The PPI network was analyzed by cytohubba tool in cytoscape, and the
   top 10 key targets were filtered by MCC method to obtain the essential
   proteins in the network. Proteins with a high score tend to be critical
   proteins.
Figure 3.
   Figure 3
   [73]Open in a new tab
   The network relationship of the first 10 key targets. The 10 key
   targets in the figure were screened by the MCC method.
   Subsequently, we screened key genes common to both using the STRING
   database for functional annotation analysis ([74]Figure 4). We found
   that the targets of action between depression and inflammation exert
   molecular functions such as regulation of cytokine activity mainly
   through cellular responses to chemical stimuli, organic substances,
   etc., based on cellular components such as extracellular gaps. The top
   20 signaling pathways obtained from KEGG signaling pathway enrichment
   analysis are shown in [75]Figure 5. In the pathogenesis of depression,
   the HIF-1 signaling pathway, PI3K-Akt signaling pathway, Th17 cell
   differentiation, hepatitis B, and inflammatory bowel disease are
   crucial in the inflammatory response. The HIF-1 signaling pathway
   indicates that there is an interdependent relationship between immune
   response and hypoxic response. Tissue hypoxia is a prominent feature of
   the inflammatory response. HIF is rapidly stabilized under hypoxia and
   is responsible for the activation of adaptive transcriptional
   responses, including the upregulation of metabolic factors such as
   vascular growth factor. In addition, studies have shown that huperzine
   activates the HIF-1α-VEGF signaling pathway in vivo, enhancing neuronal
   synaptic plasticity and thus exerting antidepressant effects ([76]20,
   [77]21). In a mouse model of depression exposed to lipopolysaccharide
   (LPS), pioglitazone with potential anti-inflammatory effects improved
   depressive behavior via upregulation of the PI3K/AKT pathway ([78]22).
   In clinical practice, various drugs, such as saffron and NGR1, have
   been shown to improve depressive behavior through the PI3K/AKT pathway
   ([79]23, [80]24). Th17 cell differentiation may, to some extent,
   reflect the risk of depression in patients. Although it has been
   clinically shown that the signature cytokine interleukin 17A (IL-17A)
   produced by Th17 cells does not significantly correlate with the
   severity of depression, Th17 cells are involved in the gut-brain axis
   to mediate the stress response, possibly by promoting
   neuroinflammation, microglia, and astrocyte activation thereby neuronal
   damage triggering depressive symptoms. Therefore, Th17 cells may be a
   promising target for the treatment of depression ([81]25–27). In
   conclusion, multiple signaling pathways are intertwined in the
   pathogenesis of depression, with the neuroimmune system signaling
   pathway playing an important role.
Figure 4.
   [82]Figure 4
   [83]Open in a new tab
   Enrichment analysis.
Figure 5.
   [84]Figure 5
   [85]Open in a new tab
   Pathway analysis.
3.2. Anti-inflammatory effect analysis of antidepressants
   Finally, we calculated the distance between 6,100 drugs and the
   depression disease module to rank them. The top 20 drug-disease
   proximity profiles are shown in [86]Table 3. We found that most of the
   FDA-approved antidepressants were ranked highly. As shown in [87]Figure
   6, we mapped the network relationships between the first three drugs
   and the depressive disorders module. The drugs shown in [88]Figures
   6B,[89]C are all FDA-approved antidepressants. We found that they have
   more acting relationships with the depressive disorder’s module in the
   network, which aligns with our expectations. Even more interesting is
   that these drugs usually also have anti-inflammatory effects, which
   suggests that depression and inflammation have potential similarities
   in terms of pathogenesis. This implies that anti-inflammatory
   strategies can alleviate depressive symptoms to some extent. In
   addition, drugs that treat other diseases and rank highly, we speculate
   that they may have a positive effect on the treatment of depression,
   although more clinical trials are needed to determine the results. We
   found that most of the FDA-approved drugs with antidepressant effects
   ranked high, and our analysis revealed that these drugs could usually
   treat not only depression but also diseases closely related to
   inflammation, such as cancer, chronic neuropathic pain, and diabetes.
   In other words, these drugs are likely to exert antidepressant effects
   by inhibiting inflammatory pathways or reducing levels of inflammatory
   factors. These drugs likely exert their antidepressant clinical effects
   by exerting anti-inflammatory effects. Thus, the mechanism by which
   antidepressants act is likely to be related to anti-inflammation, and
   there is a correlation between depression and inflammation. For
   example, among the top three drugs, Perphenazine is an antipsychotic
   phenothiazine derivative found in clinical studies to treat ear
   swelling mediated by 12-o-tetradecanoylphorbol-13-acetate (TPA) and
   oxazolone (OXA) and exert anti-inflammatory effects in addition to
   psychiatric disorders ([90]28); Clomipramine is a tricyclic
   antidepressant that has been clinically found to also reduce
   LPS-induced neuroinflammation by partially modulating NLRP3 ([91]29);
   Amitriptyline is also a tricyclic antidepressant, and many studies have
   found that Amitriptyline, in addition to its antidepressant and
   analgesic effects, can exert anti-inflammatory effects in humans and
   animal models of acute and chronic inflammation ([92]30, [93]31). In
   addition to this many of the top-ranked drugs with antidepressant
   effects have been found to have some anti-inflammatory effects in
   clinical studies, and together these further assists in validating our
   prediction that there is a correlation between depression and
   inflammation. Second, by looking at the network action relationship map
   of disease-drug key targets, we found that most of the top-ranked drugs
   treated depression well and produced more associations with disease
   proteins in the network. This suggests that our method can measure the
   effect of drugs on the disease to some extent. It also suggests that
   drugs that are ranked high but are not currently used clinically as
   depression treatment are likely to be potentially effective for the
   treatment of depression, i.e., the indications for these drugs are
   likely to be similar to depression in terms of pathogenesis. More
   clinical trials are needed in the future to test our hypothesis.
Table 3.
   Top 20 drug-disease proximity.
   Number Drug Z-value Is it an antidepressant Indication
   1 Perphenazine −9.633 No Perphenazine is a phenothiazine used to treat
   schizophrenia as well as nausea and vomiting.
   2 Clomipramine −9.249 Yes Clomipramine is a tricyclic antidepressant
   used in the treatment of obsessive–compulsive disorder and disorders
   with an obsessive–compulsive component, such as depression,
   schizophrenia, and Tourette’s disorder.
   3 Amitriptyline −9.236 Yes Amitriptyline is a tricyclic antidepressant
   indicated in the treatment of depressive illness, either endogenous or
   psychotic, and to relieve depression associated anxiety.
   4 Selegiline −9.214 Yes Selegiline is a monoamine oxidase inhibitor
   used to treat major depressive disorder and Parkinson’s.
   5 Minaprine −8.99 Yes Minaprine is a psychotropic drug that has proved
   to be effective in the treatment of various depressive states.
   6 Prasugrel −8.744 No Prasugrel is a P2Y12 platelet inhibitor used to
   reduce the risk of thrombotic cardiovascular events in unstable angina
   or non-ST-elevation myocardial infarction (NSTEMI), and in patients
   with STEMI when managed with either primary or delayed PCI.
   7 Sertraline −8.698 Yes Sertraline is a selective serotonin reuptake
   inhibitor (SSRI) indicated to treat major depressive disorder, social
   anxiety disorder, and many other psychiatric conditions.
   8 Sorafenib −8.695 No Sorafenib is a kinase inhibitor used to treat
   unresectable liver carcinoma, advanced renal carcinoma, and
   differentiated thyroid carcinoma.
   9 Tranylcypromine −8.631 Yes Tranylcypromine is a monoamine oxidase
   inhibitor used to treat major depressive disorder.
   10 Troglitazone −8.59 No For the treatment of Type II diabetes
   mellitus. It is used alone or in combination with a sulfonylurea,
   metformin, or insulin as an adjunct to diet and exercise.
   11 Fluphenazine −8.564 No Fluphenazine is a phenothiazine used to treat
   patients requiring long-term neuroleptic therapy.
   12 Duloxetine −8.563 Yes Duloxetine is a serotonin norepinephrine
   reuptake inhibitor used to treat generalized anxiety disorder,
   neuropathic pain, osteoarthritis, and stress incontinence.
   13 Efavirenz −8.521 No Efavirenz is a non-nucleoside reverse
   transcriptase inhibitor used to treat HIV infection or prevent the
   spread of HIV.
   14 Trimipramine −8.488 Yes Trimipramine is a tricyclic antidepressant
   used to treat depression.
   15 Desipramine −8.44 Yes Desipramine is a tricyclic antidepressant used
   in the treatment of depression.
   16 Meperidine −8.396 No Meperidine is an opioid agonist with analgesic
   and sedative properties used to manage severe pain.
   17 Lopinavir −8.395 No Lopinavir is an HIV-1 protease inhibitor used in
   combination with ritonavir to treat human immunodeficiency virus (HIV)
   infection.
   18 Esketamine −8.323 Yes Esketamine is a NMDA receptor antagonist used
   for treatment-resistant depression.
   19 Pipotiazine −8.243 No Pipotiazine is an antipsychotic indicated for
   the management of chronic, non agitated schizophrenic patients.
   20 Phenobarbital −8.161 No Phenobarbital is long-lasting barbiturate
   and anticonvulsant used in the treatment of all types of seizures,
   except for absent seizures.
   [94]Open in a new tab
   The shortest path between the drug and the disease protein is
   calculated, and then the Z-value is calculated after eliminating the
   bias caused by the different number of targets of different drugs; the
   smaller the Z-value, the more effective the drug is.
Figure 6.
   [95]Figure 6
   [96]Open in a new tab
   The key targeted relative maps are visualized between the top-3 drugs
   and depression. Where (A) presents the perphenazine-depression key
   target interaction map, (B) presents the clomipramine-depression key
   target interaction map, and (C) presents the amitriptyline-depression
   key target interaction map. The drugs at the top of the list were more
   closely associated with targets for depression. The undistorted images
   could be found in the [97]supplementary materials.
   Also importantly, we hope to reposition drugs by clarifying the
   inflammatory mechanisms of depression, i.e., “new use of old drugs.”
   Because drug development is often time-consuming, costly, and has a low
   success rate, new use of old drugs is a strategy to use old drugs or
   drugs in development for indications beyond the original approval and
   to expand their scope of application and use, which not only saves a
   lot of time and resources but also significantly accelerates the drug
   development process. Based on this scenario, we predict that the
   top-ranked, i.e., other drugs in the network with a similar distance to
   the depression disease module, may be useful for the treatment of
   depressed patients, and more reliable evidence will need to be provided
   by a large number of clinical trials.
4. Discussion
   Diseases often occur not because of a defect in one gene but as a
   result of coordinated interactions between different genomes. In this
   paper, we examined the co-morbidity of depression based on the human
   protein interactome network, using a network analysis approach based on
   the depression disease module. We further investigated and predicted
   the correlation between depression and inflammation by calculating the
   distance between the depression disease module and other disease
   proteins, and by combining previous clinical studies to identify
   possible shared mechanisms between co-morbidities. A genetic study
   found that endothelial dysfunction and inflammation factors were
   definitively correlated with depressive symptoms in patients with heart
   disease ([98]32). Similarly, the results of a clinical trial study
   assessing depression, metabolic syndrome, and inflammatory markers
   showed that increased appetite in depressed patients was positively
   correlated with CRP and HIF-α ([99]33). The findings of this
   immunometabolism form of depression study suggest that we must be aware
   of disease co-morbidities in clinical treatment and consider
   immunometabolism and other pathways to intervene in depressive
   episodes. The important role of neuroinflammation in the pathogenesis
   of these co-morbidities suggests that there is likely a common feature
   between depression and these disorders, namely neuroimmune mechanisms.
   This suggests that we should pay attention to common disease
   co-morbidities in clinical practice to improve the treatment outcome.
   Secondly, we screened critical targets based on the magnitude of target
   centrality, including the pro-inflammatory factors TNF-α and IL-6 and
   the anti-inflammatory factors IL-4 and IL-10. Numerous clinical studies
   have shown that depression is closely related to the concentration of
   inflammatory factors. Patients with depression usually have higher
   concentrations of pro-inflammatory factors such as TNF-α and IL-6 than
   healthy controls. As the disease progresses, their body usually has
   lower concentrations of the anti-inflammatory factor IL-10 than healthy
   controls ([100]34). Studies have found that IL-6 concentrations are
   closely associated with depression and that the pro-inflammatory factor
   IL-6 may be involved in the brain inflammatory response through
   multiple pathways. Among others, an animal study showed that Toll-like
   receptor 4 genes and cytokine receptor genes are expressed in the
   choroid plexus (CP) and that stimulation of these receptors by
   cytokines induces the synthesis and eventual secretion of IL-6 into the
   blood-cerebrospinal fluid (CSF) ([101]35). This, in turn, alters areas
   of the brain that regulate mood-related conditions and induces
   depressive symptoms. Through enrichment and pathway analysis of
   critical targets, we identified essential pathways related to hypoxia,
   inflammatory diseases, Etc. Several previous studies demonstrated the
   therapeutic effects of hypoxic preconditioning in a rat model of
   depression ([102]36). In peripheral blood cells from depressed
   patients, studies likewise found that mRNA expression of HIF-1 and its
   target genes were mainly associated with depressive symptoms ([103]37).
   In recent years, the hypoxic pathway that triggers neurodegenerative
   lesions has received attention. The hypoxia-inducible factor (HIF)
   plays a vital role in cellular biological processes and adaptation to
   cellular stress induced by hypoxic environments and is an essential
   transcriptional regulator. Therefore, we hypothesize that the HIF-1
   signaling pathway is likely to play an important role in preventing
   depression. It also can help us identify and diagnose patients with
   atypical depression symptoms at the molecular level.
   Finally, we ranked the drugs by finding a suitable algorithm to
   calculate the distance between drugs and disease modules after
   eliminating errors due to, for example, different numbers of drug
   targets. Looking at the top-ranked drugs and combining them with modern
   clinical studies, we found that most of the FDA-approved
   antidepressants also have anti-inflammatory effects, implying that the
   mechanism of action by which these drugs exert their antidepressant
   effects is likely to be the anti-inflammatory pathway at the same time.
   In an animal study, clomipramine was shown to attenuate
   lipopolysaccharide (LPS)-induced depression in a mouse model by
   partially modulating NLRP3 inflammatory vesicles ([104]29). Among them,
   NLRP3 inflammatory vesicles are present in neurons and glial cells
   ([105]38), which are involved in many critical innate immune processes,
   such as infection and inflammation. That is, NLRP3 inflammatory
   vesicles may be important in triggering depression. At the same time,
   we hypothesized that drugs with anti-inflammatory effects also have the
   potential to exert antidepressant efficacy, and the results of a
   systematic evaluation of the efficacy and safety of anti-inflammatory
   drugs in patients with major depression showed that anti-inflammatory
   drugs not only help to treat depressed patients and exert
   antidepressant effects but also are quite safe ([106]39).
   In conclusion, it is reasonable to conclude that there is a strong
   correlation between depression and inflammation, whether from the
   perspective of co-morbidity, signaling pathway analysis, or
   antidepressant drug action analysis, which provides a basis for the
   inclusion of anti-inflammatory strategies in the treatment of
   depression in the future and provides a feasible approach to explore
   the inflammatory mechanisms of depression. It also provides some
   reference ideas for new applications of old drugs to complex diseases.
   This has some applicability in both clinical treatment strategies and
   drug development.
   There are several limitations to this study. Our study is based on
   network predictions, and more clinical practice is needed to explore
   the relationship between neuroinflammation and the onset of depression
   and its specific mechanisms of action. In addition, we only focused on
   the use of the drug without refining the therapeutic effects and
   adverse effects. Therefore, further studies are necessary to address
   these issues and provide a more comprehensive evaluation of their
   efficacy and safety. Despite these limitations, our study provides
   preliminary insights into the potential role of anti-inflammatory drugs
   as a treatment for depression and merits further investigation.
Data availability statement
   The original contributions presented in the study are included in the
   article/Supplementary materials, further inquiries can be directed to
   the corresponding author.
Author contributions
   YZ: conceptualization, funding acquisition, and supervision and project
   administration. YZ, HP, and JL: methodology. HP: formal analysis. XH:
   resources. YW: suggestion. XH, HP, YZ, and YL: writing—original draft.
   XH, HP, and JL: visualization. All authors contributed to the article
   and approved the submitted version.
Funding
   This study was supported by Scientific and Technological Innovation
   Project of China Academy of Chinese Medical Sciences (No. C12021A05042,
   No. CI2021A05401), the National Natural Science Foundation of China
   (No. 81674101), National Key Technology Support Program (No.
   2012BAI25B02), Self-selected subject of China Academy of Chinese
   Medical Sciences (No. Z0217).
Conflict of interest
   The authors declare that the research was conducted in the absence of
   any commercial or financial relationships that could be construed as a
   potential conflict of interest.
Publisher’s note
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   reviewers. Any product that may be evaluated in this article, or claim
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Acknowledgments